Megatron is the founder of the Decepticon uprising, and their most well-known and feared leader. As a young, charismatic leader forged in battle and the heritage of war, he began to believe in a grand purpose for his race, it is the Transformers' glorious destiny to rule an empire which will span the universe. The opposite of his mortal enemy Optimus Prime, he feels great contempt for other Transformers who, he feels, betray their proud heritage by demanding peace and cooperation with weaker life forms.
It is the destiny of the Decepticons to bring order to the universe through conquest, though in the millions of years since coining this purpose it remains to be seen how much of his mission statement is altruistic... and how much of it is mere words built to fuel warriors to further his desire for personal power. Megatron will attempt almost anything to achieve his goals, but his schemes are rarely in any way subtle. This is perhaps due to his arrogance, but his pride is not so strong as to dissuade him from abandoning a battle he is losing. Some would question his sanity, though these few are mostly now dead by his hands...or his Fusion cannon, depending on his mood. Sometimes he uses his energon mace to strike them down. It does not matter how they die. Death by the hands of Megatron is an honor.Megatron is very powerful and utterly ruthless. There have been several interpretations of his character; some see Megatron as a strategic leader who calls the shots from afar, whilst others see him as a tactical battlefield commander who leads by brutal example. Unlike many other villains in popular fiction, Megatron was not generally depicted as overly chaotic or insane. He was highly aggressive and a megalomaniac, but there was usually a consistent rationale behind his actions, although Megatron was often the only one who could perceive this.
There have been some sparing occasions where Megatron displays a personal sense of fair play and even honor, a complexity that is most evident in his complicated relationship with Optimus Prime. There is an unspoken mutual respect between the two leaders, born of each knowing the other better than anyone else. Megatron at times seems to derive enjoyment from the perpetual conflict that exists between them — the pleasure of ending the life of Optimus Prime will be Megatron's and Megatron's alone, and to ensure this, he has aided Prime in the face of greater threats, such as the Combaticons or Jhiaxus's second generation Cybertronians. In instances such as these, the two have come to face the fact that were it not for their diametrically opposed ideology and views, in another life, the two could be comrades — a fact that Optimus Prime views as a tragedy, but which provides Megatron with amusement.
Originally, Megatron was able to transform into a Walther P38, delivering more focused energy blasts. He can shrink and reduce his mass as he transforms, assuming sizes that comfortably allow either another Transformer or even a human being to wield him. In one instance (the episode S.O.S. Dinobots ), he retained his full size and connected to jet-mode Starscream's underside. His imposing robot form is dominated by his primary weapon - his arm-mounted fusion cannon, capable of leveling a city block in one blast. He can also sub-dimensionally link the weapon to a black hole, generating even more powerful antimatter blasts. However, this ability is almost never used, as it leaves him extremely weak and vulnerable.
He has a secondary weapon barrel mounted on his back, and can retract and replace his right hand with an energy flail. He can fire electrical blasts from his hands, laser blasts from his eyes on at least one occasion (The Autobot Run) and can reprogram computers with a port in his head. He has proven to be impervious to mind control in some instances. He also wields an energon morning star on rare occasions. According to his original tech spec, Megatron has no known weaknesses. This does not, however, spare him from defeat at the hands of his enemies.
For all his famed battle prowess and tactical ability, Megatron's complacent overconfidence often causes him to overlook some vital strategic detail. Also, he has a bad habit of ordering a retreat at the first sign that the tables have turned against his side in battle (even when the Decepticons still maintain the overall strategic advantage). Another factor that could contribute to his losses is his rough relationship with some of his own troops (most specifically Starscream). Despite his lust for galactic domination, one of Megatron's key priorities remains the safety and health of Cybertron; and from his viewpoint, the best way to accomplish that is for him to conquer it.